Winter Visitor Model
This model represents species that are present only during the winter period, with a winter peak, activity spanning the year boundary, and near-absence through spring and summer.
It provides a minimal explanation for patterns seen in the seasonal analysis of observations, showing that a small number of simple processes can produce:
- Winter-centred presence
- Distinct arrival phases
- Extended absence through summer
The model does not attempt to describe detailed ecological mechanisms. Instead, it offers a way of understanding how seasonal structure and timing combine to produce the observed patterns.
Concept
This model describes species that are present during the winter months, with activity spanning the end and beginning of the year.
It answers the question:
When is the species present?
Like the seasonal model, presence is limited to part of the year. Unlike it, the active period crosses the year boundary.
The model defines a seasonal target, representing expected activity through the year. The observed signal then adjusts towards this target over time.
The target combines:
- A winter component, representing peak presence
- An optional autumn component, representing arrival
- A summer suppression, reducing activity during the off-season
Together, these produce a cycle that rises through autumn, peaks in winter, and falls away into spring.
Model Parameters
A small number of parameters control the behaviour of the model:
| Parameter | Purpose |
|---|---|
| INITIAL_Y | Sets the starting value of the modelled signal |
| BASELINE | Sets any persistent background level (typically near zero for winter visitors) |
| WINTER_WEIGHT | Controls the strength of the winter peak |
| AUTUMN_WEIGHT | Controls the strength of the autumn arrival phase |
| SUMMER_DIP | Controls the strength of the summer suppression |
| WINTER_PEAK | Sets the timing of peak winter presence |
| AUTUMN_PEAK | Sets the timing of autumn arrival |
| SUMMER_LOW | Sets the timing of lowest summer activity |
| WINTER_WIDTH | Controls how concentrated the winter peak is |
| AUTUMN_WIDTH | Controls the breadth of the arrival phase |
| SUMMER_WIDTH | Controls the breadth of the summer low |
| GROWTH_RATE | Controls how quickly activity rises towards the seasonal target |
| DECAY_RATE | Controls how quickly activity declines |
Together, these parameters define:
- When the species arrives and peaks
- How concentrated the winter period is
- How gradually or abruptly the species appears and disappears
- How strongly the species is absent through summer
All timing parameters are expressed in months on a circular 12-month scale.
Mathematical Form
The model is a first-order system:
dy/dt = rate × (target(t) - y)
Where:
- y(t) is a relative, dimensionless measure of observable activity
- target(t) is the seasonal activity target
- rate is selected using GROWTH_RATE or DECAY_RATE depending on whether the signal is rising or falling
The target function is constructed from smooth periodic components:
target(t) = winter(t) + autumn(t) - summer(t) + BASELINE
Each component is a smooth function over a 12-month cycle, allowing continuous variation without discontinuities.
Model Behaviour
When applied over a full year, the model produces a winter-centred cycle:
- Activity rises through autumn as the species arrives
- Peaks in mid-winter
- Declines through late winter and early spring
- Remains close to zero through summer
The shape depends on:
- The timing and strength of the winter and autumn components
- The breadth of the winter peak
- The strength of summer suppression
- The rate at which the system responds to change
Unlike the seasonal presence model, the season is not bounded within a single part of the calendar year. Instead, it wraps across the year boundary.
Normalisation
Model outputs are expressed as a relative measure of activity.
To allow comparison across species, results are normalised so that:
- 1.0 → peak activity
- 0.5 → half of peak activity
- 0.0 → effectively zero
This focuses attention on the timing and shape of seasonal variation.
Parameter Interpretation
After parameter fitting, the parameters are broadly interpretable as follows:
- WINTER_PEAK → timing of peak presence
- AUTUMN_PEAK → timing of arrival
- WINTER_WIDTH → concentration of winter activity
- AUTUMN_WIDTH → spread of the arrival phase
- SUMMER_DIP / SUMMER_LOW → strength and timing of absence
As with all simple models:
- Parameters should be treated as estimates rather than exact dates
- Different combinations may produce similar curves
- Interpretation is most reliable when considered alongside the fitted curve itself
In practice, each species can be described by both:
- Its fitted parameters
- The shape of its simulated seasonal curve
Together, these form a compact description of seasonal presence.
Tool
ODE Solver
A simple tool for exploring time-based models
The seasonal presence and detectability models were developed using a small, general-purpose ordinary differential equation solver, designed for experimentation and visualisation.
It allows simple systems to be defined and explored over time, making it possible to test how patterns might arise from underlying processes.
The application, the models, and instructions on how to run them are provided in the GitHub repository.